Bombus senex Vollenhoven DISTRIBUTION: Sumatran Region. Bombus supremus Morawitz DISTRIBUTION: Oriental Region. Bombus tichenkoi (Skorikov) TAXONOMIC STATUS: B. sushkini and B. saltuarius have been regarded both as conspecific (Bischoff, 1936) and as separate species (Skorikov, 1931; Tkalcu, 1974a). B. saltuarius is said to be differentiated by the morphology of the male genitalia (Skorikov, 1931), although I have seen no males of this species. More evidence is awaited. DISTRIBUTION: Palaearctic, Oriental Regions, Arctic borders. Bombus trifasciatus Smith TAXONOMIC STATUS: This species is now recognised as a separate species from gene data by Hines & Williams (2012) and by Huang et al. (2015). DISTRIBUTION: Oriental Region. Bombus ussurensis Radoszkowski DISTRIBUTION: Oriental, Japanese, Palaearctic Regions. Bombus eximius Smith DISTRIBUTION: Oriental Region. Bombus festivus Smith DISTRIBUTION: Oriental Region. Bombus friseanus Skorikov DISTRIBUTION: Oriental Region. Bombus incertus Morawitz DISTRIBUTION: Palaearctic Region. Bombus keriensis Morawitz TAXONOMIC STATUS: Several of these nominal taxa have been treated as separate species (e.g. Skorikov, 1931), although B. keriensis has also long been considered to be a broadly-distributed and variable species, including both yellow-banded and white-banded individuals throughout much of its range (Reinig, 1935, 1939; Williams, 1991 [pdf]). The taxon alagesianus is morphologically closely similar to B. keriensis. Evidence from comparisons of a few COI barcodes is inconclusive at present. Until more evidence to the contrary is available from more detailed studies of patterns of variation, I shall continue to treat them as parts of a species-complex. DISTRIBUTION: Oriental, Palaearctic Regions. Bombus ladakhensis Richards DISTRIBUTION: Oriental Region, Palaearctic border. Bombus miniatus Bingham TAXONOMIC STATUS: B. miniatus has been considered conspecific with B. pyrosoma, B. formosellus and B. friseanus (Williams, 1991 [pdf]). Evidence of intermediates between B. miniatus and B. friseanus is not strong, but perhaps not least because so little material is available from where these taxa occur in close proximity in the eastern Himalaya. The few workers and males from this area that I have seen are difficult to assign to either taxon with any confidence, although the queens are closer to the colour pattern of B. miniatus (Williams, 1991 [pdf]). From COI barcodes, these taxa appear to remain discrete and are likely to be separate species. More evidence is awaited. B. eurythorax and B. stenothorax are closely similar in morphology and colour pattern to B. miniatus. I know of no reason why these nominal taxa should not be considered conspecific. NOMENCLATURE: With Psithyrus regarded as being a subgenus of the genus Bombus, B. flavothoracicus Bingham (1897) becomes a junior secondary homonym in Bombus of Psithyrus campestris var. flavothoracicus Hoffer (1889) (deemed to be subspecific, see ICZN, 1999: Article 45.6), and therefore B. flavothoracicus Bingham is invalid (ICZN, 1999: Article 57). For this species, the oldest available name is B. miniatus, which becomes the valid name. The only subsequent publications of which I am aware that use the name B. flavothoracicus for this taxon as a species are by Tkalcu (1974b), Wang (1982) and Macior (1990), so this change of valid name is not a serious disruption of common usage. DISTRIBUTION: Oriental Region. Bombus pyrosoma Morawitz TAXONOMIC STATUS: B. pyrosoma has been considered conspecific with B. friseanus (Bischoff, 1936) and has been considered conspecific with B. formosellus, B. friseanus, and B. flavothoracicus (= B. miniatus) (Williams, 1991 [pdf]). From COI barcodes, these taxa appear to remain discrete and are likely to be separate species. More evidence is awaited. DISTRIBUTION: Oriental Region, Palaearctic border. Bombus richardsiellus (Tkalcu) DISTRIBUTION: Oriental Region. Bombus rufipes Lepeletier |